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Politics : Evolution -- Ignore unavailable to you. Want to Upgrade?


To: Brumar89 who wrote (39484)7/24/2013 3:47:49 PM
From: Brumar89  Respond to of 69300
 
The Quiet Passing of Punctuated Equilibrium, Finally!

Ray Bohlin July 22, 2013 6:11 PM | Permalink

In 1984, I was the primary co-author of a book titled The Natural Limits to Biological Change, with Lane Lester. One of my observations was that we had ample evidence of biological change in both plants and animals but there always seemed to be a limit. Only so much protein content in corn or sugar content in beets, or so many types of roses and dogs, or people three feet tall and some even eight feet tall but no-two foot dwarves or ten-foot goliaths. Of course the Darwinian evolutionary solution was new variation added through the process of mutation.

Neo-Darwinism had reigned supreme for over fifty years but there was a new kid on the block, what Stephen Gould and Niles Eldredge termed "punctuated equilibrium." In Natural Limits we critiqued both neo-Darwinism and punctuated equilibrium. Therefore I read with interest Stephen Meyer’s discussion of "Punk Eek" in Chapter 7 of Darwin’s Doubt.

From the beginning, punctuated equilibrium was primarily an observation from paleontology that most species seemed to stay the same over millions of years (stasis or equilibrium) and suddenly a new species appeared with no smooth transition (punctuation) from the previous species. Gould, Eldredge, and Stephen Stanley wrote numerous articles and books over twenty years summarizing and applying their ideas to different types of organisms in numerous geological time frames. This was all fine and good as a paleontological observation, but how was this rapid punctuation supposed to happen biologically?

Gould, in particular, was careful to point out that punctuated equilibrium was a descriptive theory of large-scale patterns over geological time, not a theory of genetic process. But if genetic process could not accomplish large-scale patterns, the observation becomes mute. One of the candidates for explaining rapid change through speciation was some kind of developmental change. At the time it was called a bifurcation. A bifurcation is defined as a series of small changes that suddenly reveals a major shift. A child’s metal cricket or frog is a good example. These toys consist of a short piece of metal, grounded at one end. As the opposite end is bent, it eventually buckles emitting the desired “click.” So perhaps with populations, mutations in a particular adaptation could accumulate over time and suddenly result in a major morphologic shift. But even in 1984 (when evolutionary developmental biology "evo-devo" was in its infancy) nearly all discussions of developmental gene mutations were discussions of lethal or near lethal changes. As we said in Natural Limits, “Mutations of genes intricately involved in development are what produce hopeless, not hopeful monsters.”

In Darwin’s Doubt, following a helpful discussion of punctuated equilibrium along with its many difficulties, Meyer quotes Gould from 2002, “I recognize that we know no mechanisms for the origin of such organismal features other than conventional natural selection at the organismic level” (p. 149). Indeed, in 1993 Gould and Eldredge published a review paper in Nature announcing that punctuated equilibrium had come of age. But towards the end of the paper under the heading of Difficulties and Prospects, they highlight the questions raised by evolutionary biologists. After dismissing a few of those objections, they admit “But continuing unhappiness, justified this time, focuses upon claims that speciation causes significant morphological change, for no validation of such a position has emerged.”

As Meyer spells out, Eldredge and Gould postulated that the speciation event itself was involved in bringing about abrupt morphological change. But as Meyer makes clear, there was no biology behind the claim, just an observation of the fossil record. Using development as the location of such change has proven extremely difficult, as Meyer also points out, especially when discussing the origin of the major body plans that appear in the Cambrian explosion. Body plans are set early in development and mutations in these early processes are universally disadvantageous. I came across this observation from an author Meyer mentions a few times, Wallace Arthur. Arthur is a population ecologist who became intensely interested in the process of major morphologic changes. His 1997 book, The Origin of Animal Body Plans: A Study in Evolutionary Developmental Biology, is a wide-ranging discussion of the problems of achieving major morphological shifts but also an attempt at a solution. Arthur addresses the major difficulty on p. 14:

“There is, however, a problem. Those genes that control early developmental processes are involved in the establishment of the basic body plan. Mutations in these genes will usually be extremely disadvantageous, and it is conceivable that they are always so” (italics in original).

But on the following page Arthur states his faith position that “The genes involved have evolved” (italics in original).

So if I may paraphrase: “Early developmental genes produce the differences in body plans, but these genes can’t be mutated without disastrous results. But we know they have evolved despite this evidence because evolution happened.”

Later, Arthur addresses the primary problem for a neo-Darwinian approach of accumulating small mutations to produce a major morphological shift (italics in original): In a developmentally explicit approach it is clear that many late changes can not accumulate to give an early one. Thus if taxonomically distant organisms differ right back to their early embryogenesis, as is often the case, the mutations involved in their evolutionary divergence did not involve the same genes as those involved in a typical speciation event, where usually the early embryogeneses of the daughter species are virtually identical. [p. 22]

Punk Eek was dead over twenty years ago but persisted on the coattails of Stephen Gould’s considerable and deserved celebrity. But with him gone, his and Elderdege’s unique contribution to evolutionary theory is finally passing quietly away.

- See more at: evolutionnews.org



To: Brumar89 who wrote (39484)7/25/2013 12:45:07 AM
From: 2MAR$  Read Replies (1) | Respond to of 69300
 
Today, geologists recognize that the rates of geological processes have varied considerably through the history of Earth and that many processes have operated in the past that may not be readily studied today.

Exactly , whats more the changing conditions in the ocean's composition with the great oxygen cycle that starts about 2.4Bil yrs ago then accummulates & builds right up to 850mil yrs ago would be a major influence for rapid change. The atmosphere goes from practically no free oxygen present to rising levels much higher approaching up into the Cambrian.

Surprise surprise, we see no discussion of this?
en.wikipedia.org



O2 build-up in the Earth's atmosphere. Red and green lines represent the range of the estimates while time is measured in billions of years ago (Ga).
Stage 1 (3.85–2.45 Ga): Practically no O2 in the atmosphere.
Stage 2 (2.45–1.85 Ga): O2 produced, but absorbed in oceans & seabed rock.
Stage 3 (1.85–0.85 Ga): O2 starts to gas out of the oceans, but is absorbed by land surfaces.
Stages 4 & 5 (0.85–present): O2 sinks filled and the gas accumulates. [1]



To: Brumar89 who wrote (39484)7/25/2013 12:53:57 AM
From: 2MAR$  Read Replies (2) | Respond to of 69300
 
Stephen "daisy chain" Meyer would be better off sticking to the "proof by beauty" or fine tuning argument, that would be more his speed. Evolution & common descent are still supported by that vast amount of gathered molecular genetic data, to suggest otherwise as he's doing here is just creationism, not even ID. (and a lie)

Poof, poof, goes the magic weasel! But nothing has been proved .



To: Brumar89 who wrote (39484)7/25/2013 12:57:08 AM
From: 2MAR$  Read Replies (1) | Respond to of 69300
 
The supernatural is always invisible, thats because its only within the limited human brain/ mind, religion is also representative of an evolutionary expression in man.

Remember how you fellows like to call "Darwinism" a religion? lol

Its gone & going far beyond that.